Nipping often means “Move, you are in my way” or “Pet me”. It is about communication. Some buns will nip if you stop petting them sooner than they find acceptable. Since bunny nipping has evolved among rabbits, they are more painful to us hooms than they would be if the nipped individual were a bun with a thick layer of protective fur.
In order to understand if this is aggressive biting or a demand for food or attention, you need to weigh in how he looks when he nips and where the nipping takes place. If its in his area, as LBJ mentions, it could be about territory (get out of my home, horrible hooman). If he feels cornered with nowhere to run, it could be about fighting when flight isn’t possible. If he comes up to you and nips “out of the blue”, I think its more about attracting your attention to his need of food, pets or treats.
Avgression in rabbits often involve raised tail, growling (like a dog, albeit tiny) and boxing, as well as biting.
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