Hi
It’s no wonder that M is terrified of a bunny that always growls at her. It sounds like there’s a few separate problems here.
How long have you had them all?
Before bunnies can live together, they must all be spayed and neutered, and they must have been prebonded and bonded. Pop over to the bonding forum to find out a bit more about the bonding process, but it will be a while before you can start bonding them since 2 still aren’t neutered.
At this stage, none of them should be allowed to touch each other at all. Try blocking off the other cages when one is out for floor time.
I don’t know how long you’ve had M, but it sounds like shes pretty new. She probably needs time to get used to her new home, and to being surrounded by other aggressive rabbits.
To bond with her yourself, you can try lying on the floor with her, and letting her come to you. Don’t pick her up or follow her about, it must all be on her terms to make her comfortable. You can try hand feeding her when she’s a bit more confident too.
Spaying makes rabbits less territorial and aggressive, but it won’t make them less scared. You have to work on building a relationship with her.